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Triceratops

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iTriceratops
Fossil range: Late Cretaceous

Triceratops skeleton at the
National Museum of Natural History.

Conservation status

Extinct (fossil)

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Scientific classification

Kingdom:	Animalia
Phylum:	Chordata
Class:	Sauropsida
Superorder:	Dinosauria
Order:	Ornithischia
Suborder:	Marginocephalia
Infraorder:	Ceratopsia
Family:	Ceratopsidae
Subfamily:	Ceratopsinae
Genus:	Triceratops Marsh, 1889

Species

See text.

Triceratops (traɪ'sɛ.ræ.tɒps) meaning 'three-horned face' (derived from the Greek tri -/τρι- meaning 'three', ceras/κέρας meaning 'horn' and -ops/ωψ meaning 'face'<ref>Liddell & Scott (1980). Greek-English Lexicon, Abridged Edition. Oxford University Press, Oxford, UK. ISBN 0-19-910207-4.</ref>) was a ceratopsid herbivorous dinosaur genus, from the Late Cretaceous Period (from around 70-65 million years ago) of what is now North America. It lived at around the same time and place as Tyrannosaurus, Ankylosaurus and another well-known ceratopsid, Torosaurus.

Although no complete skeleton has been found, it has been estimated that Triceratops was about 9 m (30 ft) long, 3 m (10 ft) tall, and weighed around 5,400 kg (12,000 lb).

1 Discoveries and species
- 1.1 How many species?
2 Paleobiology
3 Depiction in popular media
4 References
5 External links

[edit] Discoveries and species

Image:Triceratops2.png

Triceratops was discovered by John Bell Hatcher, in 1888. Its declaration as a legitimate dinosaur came when an intact skull was found. It was named by Othniel Charles Marsh in 1889 (some time earlier, however, in 1887 near Denver, Colorado, USA, he had misidentified the Triceratops as a type of bison, giving it the name Bison alticornis). The sturdy nature of the animal's skull has ensured that many examples have been preserved as fossils, allowing variations between species and individuals to be studied. Triceratops remains have subsequently been found in Montana, South Dakota, and Wyoming, in the USA and in Saskatchewan and Alberta, in Canada.

[edit] How many species?

In the first decades after Triceratops was described, various skulls were collected, which varied to a lesser or greater degree from the original Triceratops, named T. horridus by Marsh (from Latin horridus; "rough, rugose", suggesting the roughened texture of the bones, which Marsh later admitted belonged to an aged individual). Discoverers would write these up as separate species (listed below). Eventually, however, the idea that the differing skulls might be representative of individual variation within one (or two) species gained popularity. In 1986, Ostrom and Wellnhofer <ref>Ostrom, J. H., and P. Wellnhofer. 1986. The Munich specimen of Triceratops with a revision of the genus. Zitteliana 14: 111 - 158.</ref>published a paper where they proposed there was only one species - Triceratops horridus. Part of the rationale is that generally there are only one or two species of any large animal in a region (e.g. elephant or giraffe in modern Africa). A few years later, Cathy Forster reanalysed<ref>Forster CA (1996): Species resolution in Triceratops: cladistic and morphometric approaches. J.Vert.Paleont. 16(2): 259-270</ref> Triceratops material more comprehensively and concluded that the remains fell into two species, T. horridus and T. prorsus, although the distinctive skull of T. (now tentatively Diceratops) hatcheri differed enough to warrant a separate genus.

Triceratops species:

T. horridus Marsh, 1889 (type species)
T. prorsus Marsh, 1890

Image:Triceratops brussels email.jpg

Nomina dubia:

T. albertensis Sternberg, 1949
T. alticornis Marsh, 1887 (originally 'Bison')
T. eurycephalus Schlaikjer, 1935
T. galeus Marsh, 1889
T. ingens Lull, 1915
T. maximus Brown, 1933
T. sulcatus Marsh, 1890

Misassignments:

T. brevicornis Hatcher, 1905 (=T. prorsus)
T. calicornus Marsh, 1898 (=T. horridus)
T. elatus Marsh, 1891 (=T. horridus)
T. flabellatus Marsh, 1889 (=T. horridus)
T. hatcheri Lull, 1907 (=Diceratops hatcheri)
T. mortuarius Cope, 1874 (also nomen dubium; originally Polyonax; =Polyonax mortuarius)
T. obtusus Marsh, 1898 (=T. horridus)
T. serratus Marsh, 1890 (=T. horridus)
T. sylvestris Cope, 1872 (nomen dubium; originally Agathaumas; =Agathaumas sylvestris)

[edit] Paleobiology

Although Triceratops is commonly portrayed as a herding animal, there is currently no solid evidence that it lived in herds. Unlike other horned dinosaurs, some of which are known from sites preserving dozens or hundreds of individuals, all Triceratops finds known at present preserve only solitary individuals.

[edit] Dentition

Its food was plants and shrubbery and its snout consisted of a sharp beak, which would have enabled it to break up and eat very tough vegetation. This beak could also be used in self-defense, as in the case of the primitive horned dinosaur Protoceratops. Behind the beak, Triceratops had a series of teeth arranged in a shearing configuration. Triceratops teeth are among the most abundant fossils in the Late Cretaceous Period of Western North America (65 mya), suggesting that it was the dominant herbivore of the time.

[edit] Gait

Triceratops possessed a sturdy build, with robust legs and five-hoofed toes. It is estimated that Triceratops was able to run at around 24 km/h (15 mph), since its short legs meant it could not take long strides.

[edit] Posture

The posture of Triceratops has long been the subject of some debate. Originally, it was believed that the front legs of the animal had to be sprawling at angles from the thorax, in order to better bear the weight of the head. This stance can be seen in paintings by Charles Knight and Rudolf Zallinger. However, ichnological evidence in Triceratops trackways seem to show that Triceratops maintained an upright stance during normal locomotion, with the knees slightly bowed out (as in the modern rhinoceros). This does not preclude a sprawling gait for confrontations or feeding.

[edit] Horns and frill

Image:Triceratops 1.jpg

Image:Triceratops 2.jpg

The distinctive skull of Triceratops had a single horn on the snout, above the nostrils and a pair of horns approximately 1 m (3 ft, 4 in) long, above the eyes. The rear of the skull bore a relatively short bony frill. Most other frilled dinosaurs had large fenestrae in their frills, while the frill of Triceratops is noticeably solid.

A number of functions have been proposed for the frill:

Defense against carnivores, such as Tyrannosaurus
Communication between herd members
Battling rival Triceratops over status, resources or territory
Courtship display
A status symbol which reflects (or determines) the individual's status in the herd
Anchor points for the jaw muscles
Increasing body area, to regulate body temperature (see also: thermoregulation)

In 2005, a BBC documentary, The Truth About Killer Dinosaurs, tested how Triceratops might have defended itself against large predators like Tyrannosaurus. To see if Triceratops could have charged other dinosaurs, as would a modern-day rhinoceros, an artificial Triceratops skull was made and propelled into simulated Tyrannosaurus skin, at 24 km/h (15 mph). The brow horns penetrated the skin but the blunt nose horn and the beak could not and the front of the skull broke. The conclusion drawn was that it would have been impossible for Triceratops to have defended itself in this way - instead, it probably stood its ground when attacked by large predators, using its horns for goring if the predator came close enough.

A recent study <ref>Goodwin MB, Clemens WA, Horner JR, Padian K: The smallest known Triceratops skull: new observations on Ceratopsid cranial anatomy and ontogeny, Journal of Vertebrate Paleontology, 2006, 26(1):103-112</ref> of the smallest Triceratops skull, ascertained to be a juvenile, shows the frills and horns developed at a very early age, predating sexual development and thus possibly important for visual communication and species recognition.

[edit] Depiction in popular media

In popular culture, Triceratops is famously known as "three-horns" due to the three prominent horns on its head and nose, which have become almost synonymous with the dinosaur. Triceratops is also the official state fossil of South Dakota, and the official state dinosaur of Wyoming.

Many (if not most) dinosaur movies have included Triceratops. One Million Years B.C. (1966) features Triceratops in a memorable but anachronistic battle with Ceratosaurus. In Jurassic Park (1993), an immobilized Triceratops is depicted as having a mysterious illness, the role given to a Stegosaurus in the novel. In the sequel movie, The Lost World: Jurassic Park, a Triceratops is captured but later set free, causing chaos. In Jurassic Park III, a herd of Triceratops are seen from an airplane as it lands at the island.

A popular children's book is Oliver Butterworth's The Enormous Egg, where a friendly Triceratops hatches from an unusually large hen's egg.

Representations based on Triceratops featured in Japan; Triceramon, a Digimon, is based on Triceratops. In Japan's Super Sentai series, in Kyoryuu Sentai ZyuRanger, Dan's, aka TriceraRanger, powers were symbolized by the Triceratops. Later in Bakuryuu Sentai Abaranger, Yukito Sanjyou, aka AbareBlue (whose name is pun off of the Triassic period), also uses the Triceratops mecha.

Likewise in the Power Rangers series which were based off the cast of Sentai, in Mighty Morphin Power Rangers, the Triceratops was Billy Cranston's power source. In Power Rangers: Dino Thunder, the Triceratops was Ethan James's zord.

In the 1980s cartoon, Dino-Riders, the Rulons used numerous Triceratops.

The Transformers characters of Slag, Triceradon and Guiledart each turn into a Triceratops.

The short-lived series Dinozaurs had the character Dino Tricera, one of the core Dino Knights, whose top horns become the Tricera Saber of Jade. He was blue with the holes of the skeleton filled in with lime green.

The last episode of the documentary series Walking With Dinosaurs briefly featured the dinosaur as the meal of a Tyrannosaurus.

The 2005 BBC documentary The Truth About Killer Dinosaurs featured a Triceratops. It was put against a Tyrannosaurus in the first episode, with the Triceratops being the victor.

In The Land Before Time and its sequels, 'Cera' is a young Triceratops. Her father, an adult, is only shown.

In Diddy Kong Racing, a non-playable Triceratops character named Tricky was the boss of Dino Domain.

In the 1933 film King Kong, there was going to be a scene where Kong fights off a group of Triceratops, although this was never seen in the completed version. In the 2005 film a fictional Triceratops-like dinosaur called Ferructus makes a quick cameo - it is seen drinking from a lake.

The first episode of the 2006 drama documentary Prehistoric Park featured Triceratops, besieged by Tyrannosaurus. A young Triceratops named Theo is also captured for the titular park.

In one or more of Edgar Rice Burroughs' Tarzan novels, cavemen are depicted riding Triceratops, beating on their frills to make them change direction. They are also erroneously shown as being carnivorous.

The 1931 film, Creation included a mother Triceratops with her babies.