Vipera

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iVipera

Asp viper, V. aspis

Scientific classification

Kingdom:	Animalia
Phylum:	Chordata
Subphylum:	Vertebrata
Class:	Reptilia
Order:	Squamata
Suborder:	Serpentes
Family:	Viperidae
Subfamily:	Viperinae
Genus:	Vipera Laurenti, 1768

Synonyms

Vipera - Laurenti, 1768
Pelias - Merrem, 1820
Chersea - Fleming, 1822
Rhinaspis - Bonaparte, 1834
Rhinechis - Fitzinger, 1843
Echidnoides - Mauduyt, 1844
Mesocoronis - Reuss, 1927
Teleovipera - Reuss, 1927
Acridophaga - Reuss, 1927
Mesovipera - Reuss, 1927
Mesohoronis - Reuss, 1927
Mesohorinis - Reuss, 1927
Latastea - Reuss, 1929
Tzarevcsya - Reuss, 1929
Latasteopara - Reuss, 1935<ref name="McD99">McDiarmid RW, Campbell JA, Touré T. 1999. Snake Species of the World: A Taxonomic and Geographic Reference, vol. 1. Herpetologists' League. 511 pp. ISBN 1-893777-00-6 (series). ISBN 1-893777-01-4 (volume).</ref>

Common names: Palaearctic vipers,<ref name="Mal03">Mallow D, Ludwig D, Nilson G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Krieger Publishing Company, Malabar, Florida. 359 pp. ISBN 0-89464-877-2.</ref> Eurasian vipers.<ref name="SB95">Spawls S, Branch B. 1995. The Dangerous Snakes of Africa. Ralph Curtis Books. Dubai: Oriental Press. 192 pp. ISBN 0-88359-029-8.</ref>

Vipera is a genus of venomous vipers. It has a very wide range, being found from from North Africa to just withing the Arctic Circle and from the British Isles to Pacific Asia.<ref name="Mal03"/> 23 species are currently recognized.<ref name="ITIS">Vipera (TSN 202180). Integrated Taxonomic Information System. Accessed on 13 August 2006.</ref>

1 Description
2 Geographic range
3 Habitat
4 Behavior
5 Reproduction
6 Venom
7 Species
8 See also
9 Cited references
10 External links

[edit] Description

Members are usually small and more or less stoutly built. The head is distinct from the neck and covered with small scales in many species, although some have a few small plates on top. The dorsal scales are strongly keeled, the anal scale is divided and the subcaudals paired.<ref name="Mal03"/>

[edit] Geographic range

They can be found in Great Britain and nearly all of continental Europe, on some small islands of the Mediterranean (Elba, Montecristo, Sicily) and the Aegean Sea, as well as in northern Africa in Morocco, Algeria and Tunisia. It also occurs across the Arctic Circle and eastwards though northern Asia to Sakhalin Island and northern Korea. <ref name="McD99"/>

[edit] Habitat

Most species prefer cooler environments. Those found at lower latitudes tend to prefer higher altitudes and dryer, rocky habitats, while the species that occur at more northern latitudes prefer lower elevations and environments that have more vegetation and moisture.<ref name="Mal03"/>

[edit] Behavior

All species are terrestrial.<ref name="Mal03"/>

[edit] Reproduction

All members are viviparous, giving birth to live young.<ref name="Mal03"/>

[edit] Venom

Most Vipera species have venom that contains both neurotoxic and hemotoxic components. Bites vary widely in severity. Smaller, northern species, such as V. berus, have only slightly less toxic venom, but inject very little. Others, such as V. ammodytes, are capable of injecting much more with devastating results. However, bites from Vipera species are rarely as severe as those from larger Macrovipera or Daboia.<ref name="Mal03"/>

[edit] Species

Species<ref name="McD99"/>	Authority<ref name="McD99"/>	Subsp.*	Common name	Geographic range<ref name="McD99"/>
V. albicornuta	Nilson & Andrén, 1985	0	Iranian mountain viper	The Zanjan Valley and surrounding mountains in northwestern Iran.
V. albizona	Nilson, Andrén & Flärdh, 1990	0	Central Turkish mountain viper	Central Turkey.
V. ammodytes	(Linnaeus, 1758)	4	Horned viper	North-eastern Italy, southern Slovakia, western Hungry, Slovenia, Croatia, Bosnia-Herzegovina, Serbia, Montenegro, Albania, Greece (including Macedonia and Cyclades), Romania, Bulgaria, Turkey, Georgia and Syria.
V. aspis^T	(Linnaeus, 1758)	4	Asp viper	France, Andorra, northeastern Spain, extreme southwestern Germany, Switzerland, Monaco, the islands of Elba and Montecristo, Sicily, Italy, San Marino and northwestern Slovenia.
V. barani	Böhme & Joger, 1984	0	Baran's adder	Northwestern Turkey.
V. berus	(Linnaeus, 1758)	2	Common European adder	From western Europe (Great Britain, Scandinavia, France) across central (Italy, Albania, Bulgaria and northern Greece) and eastern Europe to north of the Arctic Circle, and Russia to the Pacific Ocean, Sakhalin Island, North Korea, northern Mongolia and northern China.
V. bornmuelleri	Werner, 1898	0	Bornmuellers viper	Golan Heights, southern Lebanon and Syria.
V. bulgardaghica	Nilson & Andrén, 1985	0	Bulgardagh viper	The Bulgar Dagh (Bolkar Dagi) mountains, Nigde Province, south central Anatolia, Turkey.
V. darevskii	Vedmederja, Orlov & Tuniyev, 1986	0	Darevsky's viper	The southeastern Dzavachet Mountains in Armenia and adjacent areas in Georgia.
V. dinniki	Nikolsky, 1913	0	Dinnik's viper	Russia (Great Caucasus) and Georgia (high mountain basin of the Inguri River), eastward to Azerbaijan.
V. kaznakovi	Nikolsky, 1909	0	Caucasus viper	Northeastern Turkey, Georgia and Russia (eastern Black Sea coast.
V. latastei	Bosca, 1878	1	Lataste's viper	Extreme southwestern Europe (France, Portugal and Spain) and northwestern Africa (the Mediterranean region of Morocco, Algeria and Tunisia).
V. latifii	Mertens, Darevsky & Klemmer, 1967	0	Latifi's viper	Iran: upper Lar Valley in the Elburz Mountains.
V. lotievi	Nilson et al., 1995	0	Caucasian meadow viper	The higher range of the Big Caucasus: Russia, Georgia and Azerbaijan.
V. monticola	Saint-Girons, 1954	0	Atlas mountain viper	High Atlas Mountains, Morocco.
V. nikolskii	Vedmederja, Grubant & Rudajewa, 1986	0	Nikolsky's viper	Central Ukraine.
V. palaestinae	Werner, 1938	0	Palestine viper	Syria, Jordan, Israel and Lebanon.
V. pontica	Billing, Nilson & Sattler, 1990	0	Pontic adder	Known only from the Coruh valley in Artvin Province, northeastern Turkey.
V. raddei	Boettger, 1890	0	Rock viper	Eastern Turkey, northwestern Iran, Armenia, Azerbaijan, and probably Iraq.
V. seoanei	Lataste, 1879	1	Baskian viper	Extreme southwestern France and the northern regions of Spain and Portugal.
V. ursinii	(Bonaparte, 1835)	0	Meadow viper	Southeastern France, eastern Austria (extinct), Hungary, central Italy, Croatia, Bosnia-Herzegovina, northern and northeastern Albania, Romania, northern Bulgaria, Greece, Turkey, northwestern Iran, Armenia, Azerbaijan, Georgia, Russia and across the Khazakstan, Kirgizia and eastern Uzbekistan steppes to China (Xinjiang).
V. wagneri	Nilson & Andrén, 1984	0	Ocellated mountain viper	The mountains of eastern Turkey and adjacent northwest Iran.
V. xanthina	(Gray, 1849)	0	Rock viper	Extreme northeastern Greece, the Greek islands of Simi, Kos, Kalimnos, Leros, Lipsos, Patmos, Samos, Chios and Lesbos, European Turkey, the western half of Anatolia (inland eastward to Kayseri), and islands (e.g. Chalki, Kastellórizon [Meis Adasi]) of the Turkish mainland shelf.